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Crop Nº 03Poaceae (grasses)

Wheat

Triticum

Three wild grasses, joined by two chance crosses, the second of them in a farmer's field. No bread wheat has ever grown wild.

Wheat is a family, not a single plant. Einkorn, emmer, durum, spelt and bread wheat are different species, and the later ones were made by crossing: bread wheat carries the whole genomes of three wild grasses.

People in Southwest Asia gathered wild wheat for thousands of years, then farmed it for a thousand more before the plant itself changed. Today it covers more land than any other crop.

3
wild genomes in bread wheat
42
chromosomes, against 14 in each wild grass
219.5 M
hectares harvested in 2024, the most of any crop
798 M
metric tons grown in 2024
  • zízSumerian, emmer
  • GIGSumerian, wheat
  • farLatin, the old grain of Rome; emmer or spelt, disputed
  • 麥 màiChinese, wheat, barley or both in early texts
  • hwǣteOld English, linked to the word white

Wheat is the work of the first farmers of Southwest Asia, and of the generations of farmers who kept its landraces afterwards. Much of the variety they kept is missing from modern wheat.

11 chapters · 114 sources

Numbers open the source. Their color shows how settled the claim is:Established Supported Contested

About 800,000 years ago

Three grasses

Two of wheat's three genomes had already merged, long before anyone farmed.

Wild einkorn, collected near Smyrna (now İzmir) on May 7, 1854. It is a cousin of the wheats on this page's main line, not their ancestor. Collected by Benedict Balansa; photographed by the Natural History Museum, London, Triticum monococcum subsp. aegilopoides, specimen BM000086044, collected 7 May 1854. Natural History Museum, London, Department of Botany. CC BY 4.0. Cropped to remove the scale bar and color chart.

Wheat begins with wild grasses in the hills of Southwest Asia. Each has 14 chromosomes. One of them, Triticum urartu, still grows from Lebanon to Iran.

About 800,000 years ago a grass very like it crossed with a relative of the goat grass Aegilops speltoides. The seedling kept both parents' genomes whole. It was wild emmer, with 28 chromosomes. That second parent has never been found alive. It is most probably extinct.

Wild einkorn looks much the same as T. urartu, but it is a separate species and it stayed as it was.

14 → 28 → 42
Chromosomes: wild grass, emmer, bread wheat
4.5 million years
Since the B-genome grass split from its nearest living kin
23,000 to 11,700 years ago

Gathered, ground, baked

People ate wild wheat for more than ten thousand years before they grew it.

Wild emmer is a plant of rocky slopes around the upper Jordan valley. At Ohalo II, a camp on the Sea of Galilee about 23,000 years old, wild grasses were the staple food, and wild wheat and barley made up a large part of what was left behind. Starch on a grinding stone shows grass seeds being processed there: barley, and possibly wheat.

Some researchers see more than gathering. Weeds of the kind that grow in tilled fields hint at small trial plots. Others read the same remains as wild stands, cut and carried home.

In the Black Desert of Jordan, hunter-gatherers were baking some 14,400 years ago. Charred crumbs from two fireplaces at Shubayqa 1 are bread-like, and the excavators suggest that wild einkorn went into them.

more than 90,000
Plant remains studied at Ohalo II
24 of 49
Bread-like crumbs among those examined at Shubayqa 1
11,500 to 8,500 years ago

A slow taming

Farmers sowed wild wheat for a thousand years or more before the plant changed.

Farming came before the farmed plant. In northern Syria, at Jerf el Ahmar, Dja'de and Tell Qaramel, people of the tenth and ninth millennia BCE were growing wild einkorn and other wild cereals. The plants themselves had not yet changed.

A wild ear shatters when ripe and sows itself. Harvesting favors the rare ear that holds on, and the change shows in the chaff. At Nevalı Çori in south-east Turkey, about 10,000 years ago, only 14% of einkorn chaff was of the holding kind. At Çatalhöyük, a thousand years and more later, 95% was. Whether so slow a change means that nobody was selecting on purpose is argued.

Einkorn's DNA points to the slopes of Karacadağ, a volcano in south-east Turkey. Where emmer was tamed is disputed, between that region and the southern Levant, and the oldest fields with a good share of tamed ears lie in southern Syria.

14% of 282 pieces
Einkorn chaff that held its grain, Nevalı Çori
95% of 945
The same, Çatalhöyük, over a thousand years later
about 9,800 years ago
Domestic-type emmer at Chogha Golan, Iran
Over some two thousand years

The ear, rebuilt

It stopped shattering, the grain swelled, and in some wheats the husk let go.

Domestication changed the ear in three ways. First, it stopped shattering. In einkorn a single amino-acid change in one gene, Btr1, did it. Emmer has two genomes, so it needed two broken copies. Archaeologists read the result in the chaff: a wild spikelet breaks away with a smooth scar, a farmed one with a rough, torn one.

Second, the grain swelled. It did not get longer. It got wider and deeper, and it did so before the ear stopped shattering.

Third, in some wheats the husk let go. Einkorn, emmer and spelt are hulled: threshing breaks the ear into spikelets, and the grain has to be pounded out. Durum and bread wheat thresh free, largely because of one gene, Q.

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about 10–13 g
A thousand grains: wild einkorn
27 g
A thousand grains: einkorn landraces
1 amino acid
Change in Btr1 that holds the einkorn ear together
About 8,500 years ago

The second cross

Bread wheat was never wild. It began in or beside a field.

Four wheats in one German plate of 1911. Along the top, 4 is bread wheat, 5 spelt, 6 emmer and 7 einkorn. The other ears are rye, barley, oats, millet and maize. Hoffmann's Botanischer Bilderatlas, edited by E. Dennert (Stuttgart, Schweizerbart), Familie Gräser (Gramineae), plate 17, 1911. Biodiversity Heritage Library; file from the Wikimedia Commons copy of the library's own upload. Public domain. Resized.

Somewhere near the Caspian Sea, a cultivated wheat met a weed. The weed was Tausch's goat grass, Aegilops tauschii, which grows from Turkey to China. Their offspring kept every chromosome of both: three genomes, 42 chromosomes. It was bread wheat.

The usual date is 8,500 to 9,000 years ago, though estimates run from 6,000 to 11,000. No wild bread wheat exists.

The goat grass passed on only about a quarter of its own variety. Yet a rare lineage, found today only in Georgia, left its mark too, so the cross may have happened more than once. In the ground, DNA from grain about 8,400 years old at Çatalhöyük is of the three-genome kind, and two Georgian villages held bread wheat beside goat grass by about 5900 BCE.

about 25%
Share of goat-grass variation that reached bread wheat
107,891
Genes counted in the bread wheat genome
by the ear-stalk, not the grain
How bread wheat is told from durum in the ground
6700 to 3800 BCE

West with the first farmers

Emmer and einkorn inland, naked wheat along the coast, and one wheat that vanished.

Wheat left its homeland with the people who farmed it. Villagers on Cyprus had cereals from the mainland by about 10,600 years ago. Around 6700 BCE newcomers founded farming villages on the Aegean coast of Turkey. Wheat grains from the first of them, Çukuriçi Höyük, have been dated directly.

From the plains of northern Greece, farmers moved up into the Balkans with emmer and einkorn as their staples. By 5400 to 4900 BCE the same two wheats were the main crops at Biskupice in Poland. Naked wheat kept to the Mediterranean coast. Cereals reached Britain about 4000 BCE, and Sweden soon after.

A fourth wheat traveled with them, and then vanished. Archaeologists noticed its odd chaff in 2000 and named it the new glume wheat. Ancient DNA has since tied it to Timopheev's wheat, a separate species with its own pair of genomes. It died out in Europe before the end of the first millennium BCE.

an einkorn-like wheat
In the stomach of Ötzi the Iceman, 5,300 years ago
dated, and found to be medieval
Bread wheat in the Stone Age layers at Biskupice
5300 to 1500 BCE

South and east

To the Nile, the Indus and China, slowly, and with the dates still argued.

Measuring the crop and counting the harvest, from the tomb of Menna at Thebes, about 1400 to 1350 BCE. The museum does not say which grain is shown. Emmer and barley were Egypt's staples. Charles K. Wilkinson (painted copy), after the tomb painting, Harvest Scenes, Tomb of Menna, original about 1396–1347 BCE; copy painted for the museum by 1930. The Metropolitan Museum of Art, New York (accession 30.4.44). CC0 1.0. Cropped to remove a thin black border, and resized.

South of the Levant, the oldest farmed cereals in North Africa come from the Fayum oasis of Egypt, 4650 to 4350 BCE, where emmer was kept in pits lined with basketry.

Eastward the going was slow. The Zagros mountains held the crops up more than any desert did. Mehrgarh in Pakistan, which grew wheat of south-west Asian kinds, was long said to have begun about 8000 BCE. New dates on its burials give 5200 to 4900 BCE. By 3200 to 2600 BCE wheat of the bread-wheat type grew at early Harappan sites.

Herders took it through the mountains of Central Asia. Naked wheat lay at Tasbas in Kazakhstan by 2840 to 2490 BCE, and at Chap II in Kyrgyzstan, 2,000 meters up, soon after.

China is argued over. One grain from Shandong dates to about 2562–2209 BCE. Some take it at its word. Others call it an anomaly and count wheat as secure there only from about 2100 BCE.

2562–2209 BCE
Oldest directly dated wheat in China, a single find
2462–2038 BCE
Wheat grains dated at Chap II, Kyrgyzstan
8000 → about 5200 BCE
Start of Mehrgarh: old estimate and new
3200 BCE to 100 CE

The wheat of the first cities

For three thousand years the wheat of Sumer and Egypt was emmer.

Emmer in one of the oldest written records, a clay account from southern Mesopotamia of about 3100 to 2900 BCE. The museum reads it as a hand-out of barley and emmer. Unknown scribe, southern Mesopotamia, Cuneiform tablet: administrative account concerning the distribution of barley and emmer, about 3100–2900 BCE. The Metropolitan Museum of Art, New York (accession 1988.433.2). CC0 1.0. Cropped to the tablet and resized.

The wheat of the first cities was emmer. At Mesopotamian sites it is the commonest wheat, second only to barley, and Sumerian tablets record it going into beer and groats. The scribes wrote it zíz.

In Egypt emmer was the wheat, by most accounts the only one, from the start of farming until after Alexander's conquest in 332 BCE. Then durum, which threshes free, replaced it.

Rome remembered a hulled wheat as well. Pliny called far the first food of Latium, eaten as porridge long before bread. Whether far was emmer or spelt is still argued. By Pliny's century the city's grain came by sea, a third of it, in one ancient telling, from Egypt.

zíz · GIG
Emmer and wheat in Sumerian
4
Months a year Egypt fed Rome, in Josephus
1493 to 1909

Across the oceans

A handful of seed could found a wheat belt, and the stories about it grew in the telling.

Thirty-three horses and one machine, cutting, threshing and sacking wheat near Walla Walla, Washington, in 1902. Underwood & Underwood (publisher; the photographer is not named), Evolution of sickle and flail – 33 horse team harvester, cutting, threshing and sacking wheat – Walla Walla, Washington, 1902. Library of Congress, Prints and Photographs Division (LC-USZ62-71468). Public domain. Cropped to one half of the stereograph, without its card mount and printed caption.

Wheat sailed with Columbus's second voyage, among the seeds for the new town of La Isabela. Whether it grew depends on whom you read. Columbus reported a harvest. A companion wrote that it withered.

On the mainland the first sower may have been Juan Garrido, a free Black conquistador. In 1538 he petitioned the Crown as the first man to try wheat in New Spain. The claim is his own.

Farther north, single samples founded whole wheat belts. In 1842 David Fife of Ontario grew on a few grains from a cargo out of Danzig. His Red Fife became the leading wheat of the Canadian prairie, and a parent of Marquis, which ripened a week or more sooner.

180
Grains borne by one plant, in the story told of Garrido
7–10 days
Marquis's head start over Red Fife
1917 to 1970

The short straw

A dwarf wheat from Japan doubled harvests half a world away.

Old wheats stood up to a meter and a half tall. Feed them well and they fell over.

The answer came from Japan. In 1917 breeders there crossed a short local wheat, Daruma, with an American one. A descendant was registered in 1935 as Norin 10. In 1946 an American agronomist in occupied Japan sent its seed home.

In Mexico, Norman Borlaug's team began crossing it into their wheats in 1953 or 1954. The short plants put more of their growth into grain. In 1967 Pakistan, Turkey and India bought 82,000 tons of the seed, and within five years India's harvest had risen from 12.3 to 20 million tons.

The gains were real and uneven. In irrigated Punjab they came with salt, waterlogging and poorer groundwater.

12.3 → 20 Mt
India’s wheat harvest, 1964–65 and 1970
150 → 60–110 cm
Plant height before and after Norin 10's genes
Today

Wheat today

More land than any other crop, and a fifth of the world's calories.

Stem rust on wheat. The race called Ug99, found in Uganda in 1999, can infect wheats that carry the resistance gene Sr31. Yue Jin, USDA Agricultural Research Service, Close-up of stem rust on wheat (image K11192-1), published 2004. USDA Agricultural Research Service, Image Gallery. Public domain. Resized.

Wheat covers more of the Earth than any other crop: 219.5 million hectares in 2024, hardly more than in 1961. The harvest from that land has more than tripled, to 798 million metric tons. Three countries, China, India and Russia, grow two-fifths of it.

It gives people about a fifth of their calories and of their protein, and two-thirds of the crop is eaten as food. About 95% is bread wheat. Most of the rest is durum. What sets wheat apart is gluten, the stretchy protein that holds the gas in rising dough.

The old wheats hang on. Emmer is still grown in Ethiopia, India and Yemen. And the base is narrow. Modern varieties descend from two of seven old groups of landraces, and a rust race found in Uganda in 1999 could infect about 90% of the world's varieties.

798 M
Metric tons grown in 2024
1.09 → 3.64 t/ha
World average yield, 1961 and 2024
18%
Share of the world's food calories